Petersburg (graduating in 1879 and completing his dissertation in 1883), he studied during 1884–86 in Germany under the direction of the cardiovascular physiologist Carl Ludwig (in Leipzig) and the gastrointestinal physiologist Rudolf Heidenhain (in Breslau). Petersburg, where he studied chemistry and physiology. In 1870 he abandoned his theological studies to enter the University of St. There, he attended a church school and theological seminary, where his seminary teachers impressed him by their devotion to imparting knowledge. Pavlov, the first son of a priest and the grandson of a sexton, spent his youth in Ryazan in central Russia. ![]() He was awarded the Nobel Prize for Physiology or Medicine in 1904 for his work on digestive secretions. He developed a similar conceptual approach, emphasizing the importance of conditioning, in his pioneering studies relating human behaviour to the nervous system. In a now-classic experiment, he trained a hungry dog to salivate at the sound of a metronome or buzzer, which was previously associated with the sight of food. Ivan Pavlov, in full Ivan Petrovich Pavlov, (born September 14, 1849, Ryazan, Russia-died February 27, 1936, Leningrad ), Russian physiologist known chiefly for his development of the concept of the conditioned reflex.
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